The list of families below is arranged phylogenetically in the order presented in the Treatise (1997). In 2003, Peter Jell and Jonathan Adrain presented an exhaustive listing of trilobite generic names, and also included a listing according to families. Their list of families is rather different than the arrangement provided in the 1997 Treatise volume, so I have included their family arrangement differences (and other disparities) in green text. Author citations are also provided for the Agnostida and Redlichiida according to the revised Treatise (1997), and for other orders according to Jell & Adrain 2003 and a variety of specific sources. I hope it encourages trilobite enthusiasts to study the new treatise and gain insight on the classification of this diverse and fascinating group of organisms.

ORDER AGNOSTIDA Salter, 1864   SUBORDER AGNOSTINA Salter, 1864   Superfamily Agnostoidea M'Coy, 1849    Agnostidae M'Coy, 1849      Subfamily Agnostinae      Subfamily Ammagnostinae      Subfamily Glyptagnostinae [family Glyptagnostidae]    Ptychagnostidae Kobayashi, 1939    Spinagnostidae Howell, 1935      Subfamily Spinagnostinae [into family Peronopsidae]      Subfamily Doryagnostinae [family Doryagnostidae]    Peronopsidae Westergård, 1936    Diplagnostidae Whitehouse, 1936      Subfamily Diplagnostinae      Subfamily Oidalagnostinae      Subfamily Pseudagnostinae    Clavagnostidae Howell, 1937      Subfamily Clavagnostinae      Subfamily Aspidagnostinae    Metagnostidae Jaekel, 1909   Superfamily Uncertain    Phalacromidae Hawle & Corda, 1847 [into Agnostidae]    Sphaeragnostidae Kobayashi, 1939 [into Peronopsidae]   Superfamily Condylopygoidea Raymond, 1913    Condylopygidae Raymond, 1913  SUBORDER EODISCINA Kobayashi, 1939   Superfamily Eodiscoidea Raymond, 1913    Tsunyidiscidae S. Zhang, 1980    Hebediscidae Kobayashi, 1944    Calodiscidae Kobayashi, 1943    Weymouthiidae Kobayashi, 1943    Yukoniidae S. Zhang, 1980    Eodiscidae Raymond, 1913 ORDER LICHIDA Moore 1959   SUBORDER LICHINA   Superfamily Lichoidea    Lichidae    Lichakephalidae (including Eoacidaspidae?) .......... ORDER ODONTOPLEURIDA Whittington 1959  SUBORDER ODONTOPLEURINA   Superfamily Odontopleuroidea    Odontopleuridae (including Eoacidaspidae?)    Selenopeltidae [into Odontopleuridae]   Superfamily Dameselloidea    Damesellidae

ORDER REDLICHIIDA Richter, 1932   SUBORDER OLENELLINA Walcott, 1890b   Superfamily Olenelloidea Walcott, 1890    Olenellidae Walcott, 1890      Subfamily Olenellinae      Subfamily Biceratopsinae      Subfamily Bristoliinae      Subfamily Gabriellinae      Subfamily Laudoniinae      Subfamily Wanneriinae    Holmiidae Hupé, 1953      Subfamily Holmiinae      Subfamily Callaviinae   Superfamily Fallotaspidoidea Hupé, 1953    Fallotaspididae Hupé, 1953      Subfamily Fallotaspidinae      Subfamily Daguinaspidinae    Archaeaspididae Repina, 1979    Judomiidae Repina, 1979    Neltneriidae Hupé, 1953    Nevadiidae Hupé, 1953   SUBORDER REDLICHIINA Richter, 1932   Superfamily Emuelloidea Pocock, 1970    Emuellidae Pocock, 1970    Megapharanaspidae Paterson & Jago 2006   Superfamily Redlichioidea Poulsen, 1927    Redlichiidae Poulsen, 1927      Subfamily Redlichiinae      Subfamily Metaredlichiinae      Subfamily Neoredlichiinae      Subfamily Pararedlichiinae      Subfamily Wutingaspinae    Dolerolenidae Kobayashi, 1951      Subfamily Doleroleninae      ?Subfamily Paramalungiinae    Yinitidae Hupé, 1953    Mayiellidae Chang, 1966    Gigantopygidae Harrington, 1959      Subfamily Gigantopyginae      Subfamily Yiliangellinae    Saukiandidae Hupé, 1953      Subfamily Saukiandinae      Subfamily Despujolsiinae      Subfamily Resseropinae    Metadoxididae Whitehouse, 1939    Abadiellidae Hupé, 1953    Kueichowiidae Lu, 1965    Menneraspididae Pokrovskaya, 1959    Redlichinidae Zhang & Lin, 1980    Chengkouaspidae Zhang & Lin, 1980   Superfamily Paradoxidoidea Hawle & Corda, 1847    Paradoxididae Hawle & Corda, 1847    Centropleuridae Angelin, 1854    Xystriduridae Öpik, 1975

TRILOBITE ORDERS, SUBORDERS, AND SUPERFAMILIES

	ORDER	SUBORDERS	SUPERFAMILIES
	AGNOSTIDA LCam - UOrd	AGNOSTINA LCam - UOrd   EODISCINA LCam - MCam	Agnostoidea LCam - UOrd Condylopygoidea LCam - MCam Eodiscoidea LCam - MCam
	REDLICHIIDA LCam - MCam	OLENELLINA  LCam only   REDLICHIINA  LCam - MCam	Olenelloidea LCam only Fallotaspidoidea LCam only Emuelloidea LCam only Redlichioidea LCam - MCam Paradoxidoidea LCam - MCam
	CORYNEXOCHIDA LCam - UDev (Frasnian)	CORYNEXOCHINA LCam - UCam ILLAENINA UCam - UDev (Frasnian) LEIOSTEGIINA MCam - LOrd	Corynexochoidea LCam - UCam Illaenoidea UCam - UDev (Frasnian) Leiostegioidea MCam - LOrd
	LICHIDA MCam - UDev (Frasnian)	LICHINA MCam - UDev (Frasnian)	Lichoidea MCam - MDev?
	ODONTOPLEURIDA MCam - UDev (Frasnian)	ODONTOPLEURINA MCam - UDev (Frasnian)	Odontopleuroidea LOrd? - UDev (Frasnian) Dameselloidea MCam - UCam?
	PHACOPIDA LOrd - UDev (Fammenian)	PHACOPINA LOrd - UDev (Fammenian) CALYMENINA LOrd - UDev (Frasnian) CHEIRURINA LOrd - UDev (Frasnian)	Phacopoidea Dalmanitoidea Acastoidea Calymenoidea LOrd - UDev (Frasnian) Cheiruroidea LOrd - UDev (Frasnian)
	PTYCHOPARIIDA LCam - UOrd	PTYCHOPARIINA LCam - MOrd     OLENINA UCam - UOrd	Ellipsocephaloidea LCam - MCam Ptychoparioidea LCam - MOrd Olenoidea UCam - UOrd
	ASAPHIDA MCam - USil	ASAPHINA MCam - USil	Anomocaroidea MCam - UCam Asaphoidea LOrd - UOrd Dikelokephaloidea UCam only Remopleuridoidea UCam? - UOrd Cyclopygoidea LOrd - UOrd Trinucleioidea UCam - USil
	PROETIDA UCam- UPer	PROETINA UCam- UPer	Aulacopleuroidea LOrd - UPer Proetoidea LOrd - UPer Bathyuroidea UCam - MOrd?
	HARPETIDA UCam - UDev (Frasnian)	HARPETINA UCam - UDev (Frasnian)	Harpetioidea UCam - UDev (Frasnian)

Bergstrom, Jan. 1973. Organization, life, and systematics of trilobites. Fossils and Strata 2:1-69, figs. 1-16, pl. 1-5.

Chatterton, B.D.E., and S.E. Speyer. 1990. Applications of the study of trilobite ontogeny. Short Courses in Paleontology, Paleontological Society 3:116-36.

Clarkson, E. N. K. 1975. The evolution of the eye in trilobites. Fossils and Strata 4:7-31.

Clarkson, E. N. K. & R. Levi-Setti. 1975. Trilobite eyes and the optics of Descartes and Huygens. Nature 254 (1975): 663-667.

Fortey, R.A. 1990a. Ontogeny, hypostome attachment, and trilobite classification. Palaeontology 33:529-76, figs. 1-19, pl. 1.

Fortey, R.A. 1990b. Trilobite evolution and systematics. Short Courses in Paleontology, Paleontological Society 3:44-65.

Fortey, R.A. 1997. Classification. In Kaesler, R. L., ed. Treatise on Invertebrate Paleontology, Part O, Arthropoda 1, Trilobita, revised. Volume 1: Introduction, Order Agnostida, Order Redlichiida. xxiv + 530 pp., 309 figs. The Geological Society of America, Inc. & The University of Kansas. Boulder, Colorado & Lawrence, Kansas.

Fortey, R.A. 2001. Trilobite systematics: the last 75 years. J. Paleontology. 75(6):1141-51.

Fortey, R.A., and N.C. Hughes. 1998. Brood pouches in trilobites. J. Paleontology. 72(4):638-49.

Fortey, R. A. & R. M. Owens. 1999. The Trilobite Exoskeleton. in: Functional Morphology of the Invertebrate Skeleton. John Wiley & Sons.

Fortey, R. A. & R. M. Owens. 1999. Feeding habits in trilobites. Palaeontology 42(3):429-65.

Jell, P. A. 2003. Phylogeny of Early Cambrian trilobites. Special Papers in Palaeontology 70:45-57.

Jell, P.A. & J.M. Adrain. 2003  Available generic names for trilobites. Memoirs of the Queensland Museum 48(2):331-553

Jensen, S. 1990. Predation by early Cambrian trilobites on infaunal worms -- evidence from the Swedish Mickwitzia Sandstone. Lethaia 23:29-42.

Kaesler, R. L., ed. 1997. Treatise on Invertebrate Paleontology, Part O, Arthropoda 1, Trilobita, revised. Volume 1: Introduction, Order Agnostida, Order Redlichiida. xxiv + 530 pp., 309 figs. The Geological Society of America, Inc. & The University of Kansas. Boulder, Colorado & Lawrence, Kansas.

Levi-Setti, R. 1993. Trilobites (2nd Ed.). University of Chicago Press.

McNamara, K.J., and D.M. Rudkin. 1984. Techniques of trilobite exuviation. Lethaia 17:153-73.

Moore, R.C., ed. 1959. Treatise on Invertebrate Paleontology, Part O, Arthropoda 1. Geological Society of America & University of Kansas Press. Lawrence, Kansas & Boulder, Colorado. xix + 560 pp., 415 figs.

Stockton, W.L., and Cowen, R. 1976. Stereoscopic vision in one eye: paleophysiology of the schizochroal eye in trilobites. Paleobiology 2:304-15.

Whittington, H.B. 1988a. Hypostomes of post-Cambrian trilobites. New Mexico Bureau of Mines and Mineral Resource, Memoir 44:321-39, 26 figs.

Whittington, H.B. 1988b. Hypostomes and ventral cephalic sutures in Cambrian trilobites. Palaeontology 31:577-610.

Zhang, W.T. 1980. On the Miomera and Polymera (Trilobita). Scientia Sinica 23:223-34.

Zhang, Xiguang & E. N. K. Clarkson. 1990. The eyes of lower Cambrian eodiscid trilobites. Palaeontology 33:911-932.

Hawle, Ignaz & A.J.C. Corda. 1847. Prodrom einer Monographie der böhmischen Trilobiten. Abhandlungen Koeniglichen Boehmischen Gesellschaft der Wissenschaften. J.G. Calve, Prague. 176 p., 7 pl. [Family Phalacromidae]

Howell, B.F. 1935. New Middle Cambrian agnostian trilobites from Vermont. Journal of Paleontology. 9:218-21, pl. 22. [Family Spinagnostidae]

Howell, B.F. 1937. Cambrian Centropleura vermontensis fauna of northwestern Vermont. Geological Society of America Bulletin 48:1147-210, 6 pl. [Family Clavagnostidae]

Jaekel, O. 1909. Uber die Agnostiden. Zeitschrift der Deutschen Geologischen Gesellschaft 61:380-401. 23 fig. [Family Metagnostidae]

Kobayashi, T. 1939. On the Agnostids (part 1). Journal of the Faculty of Science, Imperial University of Tokyo, Section II, Geology, Minerology, Geography, Seismology. 5(5):69-198. [Suborder Eodiscina; Families Ptychagnostidae, Sphaeragnostidae]

Kobayashi, T. 1943. Brief notes on the eodiscids 1, their classification with a description of a new species and a new variety. Proceedings of the Imperial Academy, Tokyo. 19:37-42, fig. 1-5. [Families Calodiscidae, Weymouthiidae]

Kobayashi, T. 1944. On the eodiscids. Journal of the Faculty of Science, Imperial University of Tokyo, Section II. 7(1):1-74, pl. 1-2, table 1-4. [Family Hebediscidae]

M'Coy, F. 1849. On the classification of some British fossil Crustacea with notices of some new forms in the University collection at Cambridge. Annals and Magazine of Natural History (series 2) 4:161-79, 330-5, 392-414. [Superfamily Agnostoidea, Family Agnostidae]

Raymond, P.E. 1913. On the genera of the Eodiscidae. The Ottawa Naturalist 27:101-6. [Superfamilies Condylopygoidea, Eodisoidea; Families Condylopygidae, Eodiscidae]

Salter, J.W. 1864. A Monograph of British Trilobites, Part 1. Palaeontographical Society, London, Monograph, vol. for 1862. p. 1-80, pl. 1-6. [Order Agnostida, Suborder Agnostina]

Westergård, A.H. 1936. Paradoxides oelandicus beds of Oland: With the account of a diamond boring through the Cambrian at Mossberga. Sveriges Geologiska Undersökning, Series C. no. 394, Årsbok 30, no. 1:1-66, 12 figs., 12 pl. [Family Peronopsidae]

Whitehouse, F.W. 1936. The Cambrian faunas of north-Eastern Australia. Part 1: Stratigraphical outline; Part 2: Trilobita (Miomera), Memoirs of the Queensland Museum 11(1):59-112, fig. 1-4, pl. 8-10. [Family Diplagnostidae]

S. Zhang, 1980. in Zhang, Wentang, Yanhao Lu, Zaoling Zhu, Yiyuan Qian, Huanling Lin, Zhiyi Zhou, Sengui Zhang, & Jinliang Yuan. 1980. Cambrian trilobite faunas of southwestern China. Paleontologia Sinica 159 (new series B, no. 16):1-497, 105 fig., 134 pl. [Families Yukoniidae, Tsunydiscidae]

Order REDLICHIIDA:
last revised January 17, 2002

Angelin, N.P. 1854 in Angelin, N.P. 1851-1878. Paleontologia Scandinavica. Pars 1. Curstacea Formationis Transitionis [fasc. 2 (1854): p. i-ix, 21-92, pl. 25-41.] [Family Centropleuridae]

Chang, W. [also Zhang, Wentang] 1966. On the classification of Redlichiacea, with the description of new families and new genera. Acta Paleontologica Sinica 14(2):135-84, fig. 1-17, pl. 1-6. [Family Mayiellidae]

Harrington, H.J. 1959. Family Gigantopygidae. in: Moore, R.C., ed. 1959. Treatise on Invertebrate Paleontology, Part O, Arthropoda 1. Geological Society of America & University of Kansas Press. Lawrence, Kansas & Boulder, Colorado. xix + 560 pp., 415 figs. [Family Gigantopygidae]

Hawle, Ignaz & A.J.C. Corda. 1847. Prodrom einer Monographie der böhmischen Trilobiten. Abhandlungen Koeniglichen Boehmischen Gesellschaft der Wissenschaften. J.G. Calve, Prague. 176 p., 7 pl. [Family Paradoxididae]

Hupé, P. 1953. Contribution à l'étude du Cambrien inférieur et du Précambrian III de l'Anti-Atlas Marocain. Direction de la Production Industrielle et des Mines, Division des Mines et de la Géologie, Service Géologique, Notes et Mémoires 103:1-402, fig. 1-99, pl. 1-24. [Superfamily Fallotaspidoidea; Families Holmiidae, Fallotaspididae, Neltneriidae, Nevadiidae, Yinitidae, Saukiandidae, Abadiellidae]

Kobayashi, T. 1951. in Kobayashi, T. & F. Kato. 1951. On the ontogeny and ventral morphology of Redlichia chinensis, with description of Alutella nakamurai n. gen., n. sp. Journal of the Faculty of Science, Imperial University of Tokyo, Section II 8(3):99-143, 9 fig., pl. 1-5. [Family Dolerolenidae]

Lu, Y. 1965. Family Kueichowiidae. in: Lu Yenhao, Wentang Chang, Zhaoling Zhu, Yiyuan Qian & Liwen Xiang. 1965. Fossils of Each Group of China: Chinese Trilobites. Science Press, Beijing. 766 p., 135 pl. (2 vol.) In Chinese. [Family Kueichowiidae]

Öpik, A.A. 1975. Cymbric Vale fauna of New South Wales and Early Cambrian Biostratigraphy. Australian Government Publishing Service, Bureau of Mineral Resources, Geology and Geophysics, Bulletin 159:1-78, 14 fig., 7 pl. [Family Xystriduridae]

Pocock, K.J. 1970. The Emuellidae: a new family of trilobites from the Lower Cambrian of South Australia. Palaeontology 13:522-62, fig. 1-9, pl. 106-10. [Superfamily Emuelloidea, Family Emuellidae]

Pokrovskaya, N.V. 1959. Trilobitovaya fauna i Stratigraphiya Kembrijskikh otlochenij Tuvy [Trilobite fauna and stratigraphy of Cambrian deposits from Tuva]. Akademia Nauk SSSR, Trudy Geologicheskogo Instituta 27:1-199, pl. 1-11.  [Family Menneraspididae]

Poulsen, C. 1927. The Cambrian, Ozarkian and Canadian faunas of northwest Greenland. Meddelelser om Grønland 70:233-343, pl. 14-21.  [Superfamily Redlichioidea, Family Redlichiidae]

Repina, L.N. 1979. Zavisimost morfologicheskikh priznakov ot uslovii obitaniia trilobitov i otsenka ikh znacheniia dlia sistematiki nadsemeistva Olenelloidea [The dependence of morphological features upon the living conditions of trilobites and an evaluation of their importance in the classification of the Superfamily Olenelloidea]. Akademiia Nauk SSSR, Sibirskoe Ordelenie, Trudy Instituta Geologii i Geofizikii 431:11-30, 13 fig., pl. 1-2. [Families Archaeaspididae, Judomiidae]

Richter, R. 1932. Crustacea (Paläotologie) in: R. Dittler. G Joos, E. Korschelt, G. Linek, F, Oltmanns, K. Schaum, eds., Handwörterbuch der Naturwissenschaften, 2nd ed. Gustav Fisher. Jena. p. 840-64, fig.A, 1-64.  [Order Redlichiida, Suborder Redlichiina]

Walcott C.D. 1890. The fauna of the Lower Cambrian or Olenellus zone. In: 10th Annual Report of the Director, 1888-1889, United States Geological Survey. p. 509-774, fig. 44-69, pl. 43-98.  [Suborder Olenellina, Superfamily Olenelloidea, Family Olenellidae]

Whitehouse, F.W. 1939. The Cambrian faunas of north-Eastern Australia. Part 3: The polymerid trilobites (with supplement 1). Memoirs of the Queensland Museum 11(3):179-282, unnumbered figs., pl. 19-25. [Family Metadoxididae]

Zhang, W. & H. Lin, 1980. [Families Redlichinidae, Chengkouaspidae] in: Zhang, W., Y. Lu, Z. Zhu, Y. Qian, H. Lin, Z. Zhou, S. Zhang, & J. Yuan. 1980. Cambrian trilobite faunas of southwestern China. Palaeontologica Sinica 159 (new series B, no. 16):1-497, 105 fig., 134 pl. 

CORYNEXOCHIDA:
Kobayashi, T. 1935. The Cambro-Ordovician formations and faunas of south Chosen. Paleontology, Part 3: Cambrian faunas of south Chosen with a special study on the Cambrian trilobite genera and families. Journal of the Faculty of Science, Imperial University of Tokyo, Section II. 4(2):49-344, fig. 1-33, pl. 1-24.

LICHIDA & ODONTOPLEURIDA:
Moore, R.C. 1959. Order Lichida. in: Moore, R.C., ed. 1959. Treatise on Invertebrate Paleontology, Part O, Arthropoda 1. Geological Society of America & University of Kansas Press. Lawrence, Kansas & Boulder, Colorado. xix + 560 pp., 415 figs.

Pollit, J.R., R.A. Fortey & M.A Wills. 2005. Systematics of the Trilobite families Lichidae Hawle & Corda, 1847 and Lichakephalidae Tripp, 1957: The application of Bayesian inference to morphological data. J. Syst. Pal. 3(3):225-41.

Thomas A.T. and D.J. Holloway. 1988. Classification and phylogeny of the trilobite order Lichida. Phil. Trans. Royal Soc. London 321:179-262. 

Whittington, H.B. 2002. Lichidae (Trilobita): Morphology and classification. J. Paleontology 76(2):306-20.

PHACOPIDA:
Salter, J.W. 1864. A Monograph of British Trilobites, Part 1. Palaeontographical Society, London, Monograph, vol. for 1862. p. 1-80, pl. 1-6.

PTYCHOPARIIDA:
Swinnerton, H.H. 1915. Suggestions for a revised classification of trilobites. Geological Magazine (new series) 6(2):487-96, 538-45.

ASAPHIDA:
Fortey, R.A. & B.D.E. Chatterton. 1988. Classification of the trilobite suborder Asaphina. Palaeontology 31:165-222, fig. 1-27, pl. 17-19.

PROETIDA:
Fortey, R.A. & R.M. Owens. 1975. Proetida: a new order of trilobites. Fossils and Strata 4:227-39.

HARPETIDA:
Ebach, M.C. & K.J. McNamara. 2002. A systematic revision of the family Harpetidae (Trilobita). Records of the Western Australian Museum 21: 235-67.

TRILOBITA:
Walch, J.E.I. 1771. Die Naturgeschichte der versteinerungen zur erläuterung der Knorrischen Sammlung von Merkwurdigkeiten der Natur. Nürnberg.

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