Trilobites of the Wheeler Shale, Utah

Locality: House Range & Drum Mountains, Western Utah, USA
Stratigraphy: Wheeler Shale Formation
Age: Cambrian - ca 505 mya
 
The U-Dig Quarry, west of Delta, Utah.
 
 
Location of Utah today
Locality of Utah during the Early Cambrian

Western Utah is one of the best-known Cambrian fossil localities in the world. The Wheeler Shale and Marjum Formation, strata of Middle Cambrian age, exhibit various exposures throughout the House Range and nearby mountain ranges west of the town of Delta, Utah. The Wheeler Shale is named for a major feature in the House Range, the Wheeler Amphitheater. The Wheeler Shale contains interbeds of shaley limestone, mudstone, and thin platy limestone. Much of the Wheeler Shale is not particularly fossiliferous, but certain layers contain abundant trilobites and other shelly fossils. The Wheeler Shale also is known for a diverse biota of soft-bodied fossils, including many of the same taxa found in the Burgess Shale.

In the Cambrian, the continent of Laurentia (now the majority of North America), was equatorial, and oriented about ninety degrees from its current position. Close to the shorelines of Laurentia, limestone was deposited as shallow-ater reefs. Beyond the limestone belt, fine sediments built in deeper offshore contours, sometimes rapidly via undersea landslides off the reef platform. These offshore deposits along the paleoequator include much of the Wheeler Shale, the Burgess Shale of western Canada, and other sites from California through Utah to the Northwest territories of Canada. All of these sites yield remarkably preserved Cambrian fossils.

The most famous Wheeler Shale fossil is the trilobite Elrathia kingi; so common at some sites that specimens are commercially quarried and are made into novelty accessories, as well as sold to collectors and institutions all over the world. However, Elrathia is just one of about fifteen trilobite genera of the Wheeler Shale. Bathyuriscus fimbriatus is also relatively common at certain sites. Even more abundant are several species of agnostid trilobites, such a Peronopsis interstricta. These are typically less than a centimeter in length. Here are eight representative species of the Wheeler Shale:


Elrathia kingi
PTYCHOPARIIDA
Family Alokistocaridae

Peronopsis interstricta
AGNOSTIDA
Family Peronopsidae
 
Modocia typicalis

PTYCHOPARIIDA
Family Marjumiidae
 
Asaphiscus wheeleri
PTYCHOPARIIDA
Family Asaphiscidae

Bolaspidella housensis

PTYCHOPARIIDA
Family Menomoniidae

Jenkinsonia varga
PTYCHOPARIIDA
Family Alokistocaridae

Modocia laevinucha
PTYCHOPARIIDA
Family Marjumiidae

Bathyuriscus fimbriatus
CORYNEXOCHIDA
Family Dolichometopidae

It is notable that the trilobite fauna of the Wheeler Shale, being a Middle Cambrian locality, is dominated by Ptychopariida, Corynexochida, and Agnostida. In addition to trilobites, there were other species of arachnomorph (trilobite-like clade) arthropods such as Naraoia. These trilobite-like arthropods demonstrate that the group from which trilobites arose was itself successful and diverse, though being uncalcified, are only preserved under exceptional conditions, such as at exceptional lagerstätten such as the Burgess Shale and Chengjiang. Because the Burgess Shale was the first lagerstätte with such exceptional preservation, other sites with similar preservation are referred to as "Burgess Shale type" lagerstätten. This level of preservation occurs only infrequently in the Wheeler Shale.

TRILOBITA OF THE WHEELER FORMATION, UTAH

courtesy of the Western Trilobites Association
Taxa held in common in red
HOUSE RANGE
DRUM MOUNTAINS
Order Ptychopariida

Altiocculus harrisi

Alokistocare sp.

Asaphiscus wheeleri

Bathyocos housensis

Bolaspidella housensis

Bolaspidella wellsvillensis

Brachyaspidion microps

Brachyaspidion sulcatum
Elrathia kingi

Elrathina(=Ptychoparella) wheeleri

Jenkinsonia varga

Modocia brevispina

Modocia laevinucha

Modocia typicalis

Ptychoparella wheeleri


Order Agnostida

Baltagnostus eurypyx

Hypagnostus parvifrons

Peronopsis amplaxis

Peronopsis bidens

Peronopsis fallax

Peronopsis gaspensis

Peronopsis intermedius

Peronopsis interstrictus

Peronopsis montis

Peronopsis segmenta

Ptychagnostus atavus

Ptychagnostus gibbus

Ptychagnostus germanus

Ptychagnostus occultatus

Ptychagnostus seminula


Order Corynexochida

Bathyuriscus fimbriatus

Olenoides expansus

Olenoides nevadensis

Zacanthoides sp.

Order Ptychopariida
Altiocculus sp.

Asaphiscus wheeleri

Bathyuriscus sp.

Bolaspidella drumensis

Bolaspidella sp.

Elrathia kingi

Elrathia sp.

Ptychoparella sp.

Spencella sp.


Order Agnostida
Peronopsis amplaxis

Peronopsis fallax

Peronopsis gaspensis

Peronopsis interstricta

Peronopsis montis

Peronopsis segmenta

Ptychagnostus atavus

Ptychagnostus gibbus

Ptychagnostus intermedius

Ptychagnostus seminula


Order Corynexochida

Olenoides expansus

Olenoides serratus

Tonkinella breviceps

Zacanthoides divergens

Zacanthoides sp.


Order Asaphida
Glyphaspis concavus



Some Wheeler Shale literature:

Gunther, L. F., Gunther, V. G., and Gunther, G., 1994, Some Middle Cambrian fossils of Utah, in Special issue on Utah: Utah Geological Survey Public Information Series 26, p. 59-62.


Robison, Richard Ashby; 1962.
Late Middle Cambrian Faunas from the Wheeler and Marjum Formations of Western Utah. PhD Thesis, University of Texas at Austin

 

Robison, Richard Ashby; 1971. Additional Middle Cambrian Trilobites From the Wheeler Shale of Utah. Journal of Paleontology 45(5):796-804

Sundberg, Frederick A.; 1994. Corynexochida and Ptychopariida (Trilobita,Arthropoda) of the Ehmaniella Biozone (Middle Cambrian), Utah and Nevada. Natural History Museum of Los Angeles County - Contributions in Science, No.446

Utah's Cambrian Life: Evolution and Biogeography of Burgess Shale Type Fossils


Western Trilobites Association

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