Suborder Phacopina
Cephalon: proparian sutures (sometimes fused), schizochroal
eyes; glabella expands forwards; librigena typically yoked as single piece;
hypostome conterminant to impendent, some with no rostral plate; some with
genal spines.
Thorax: 10 to (typically) 11 segments, pleurae furrowed, articulating
facets distinct, rounded, angular, or spinose tips.
Pygidium: typically smaller than cephalon (but subisopygous
in Dalmanitoidea and Acastoidea), smooth or spinose.
Superfamilies: Phacopoidea, Dalmanitoidea, Acastoidea
Characterizations of the three Phacopine superfamilies:
Superfamily Phacopoidea
Cephalon: generally with strongly divergent axial furrows (glabella
greatly expanding anteriorly), anterior glabellar lobes fused into single
anterior tri-composite lobe, frontal area generally lacking (obliterated
by large glabella), eyes (when present) typically anterior; genal angle
typically rounded, without genal spines (exceptions among Pterygometopidae),
vincular furrow generally present.
Thorax: variable, but often with rounded tips.
Pygidium: typically micropygous (Phacopidae) to subisopygous
(some Pterygometopidae), not spinose.
Families: Phacopidae, Pterygometopidae
Genera: Ductina, Kainops, Phacops, Pterygometopus,
Reedops,
Superfamily Dalmanitoidea
Cephalon: typically bearing genal spines, generally with well-developed
preglabellar field, median region of glabella bearing
muscle scars; eyes tend to mesial or posterior, true vincular furrows
lacking.
Thorax: variable, but often with spinose tips.
Pygidium: typically subisopygous, often spinose (terminal or
marginal spines); axial rings of consistent depth
along length of pygidial axis, axial furrows with simple and consistent
convergence angle.
Families: Dalmanitidae, Diaphanometopidae, Prosopiscidae
Genera: Dalmanites, Diaphanometopus, Huntoniatonia, Odontocephalus,
Odontochile, Prosopiscus
Superfamily Acastoidea
Cephalon: axial furrows slightly to moderately divergent, anterior
glabellar lobes and furrows usually not fused (although furrows may be
indistinct), eyes typically distant from posterior border furrows, but
not strongly anterior; frontal glabellar lobe auxilliary
impression system triangular in outline, maximum width of glabella adjacent
to preglabellar furrow, median region of glabella devoid of muscle scars;
cephalic margin "shouldered," laterally convex course of genal margin topographically
distinct from curvature of axial margin
Thorax: pleural endings blunt, rounded, or angular, sometimes
spinose.
Pygidium: micropygous to subisopygous, sometimes spinose (expressed
as
marginal spines), pygidial axis with deep, apodemal
anterior ring furrows, with abrupt transition to shallow posterior ring
furrows; coincident decrease in angle of axial furrow convergence.
Other: often similar in general body form to Phacopoidea via
convergent evolution, but glabellar and other differences as noted above.
Families: Acastidae, Calmoniidae.
Genera: Acaste, Acastoides, Asteropyge, Comura, Calmonia,
Greenops, Metacanthina, Metacryphaeus, Pennaia, Phacopina, Psychopyge,
Treveropyge
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